Robber Fly - Laphria flavicollis
Family Asilidae - Robber Flies

Live robber flies photographed at northern Illinois. This female measures 18mm.

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The Asilidae genus Laphria contains some very convincing bumblebee mimics. As true flies, these robbers only have one set of flying wings. The second set present in all other insects, has evolved in the flies into small ball-like organs called halteres (visible as the whitish ball just under the base of the wing). Their function has not been fully explained, but it is thought they provide kinesthetic (balance and body postion) information during flight.

There are over 7,000 species of robber flies world wide; nearly 1,000 in North America.  All robber flies have stout, spiny legs, a dense moustache of bristles on the face (mystax), and 3 simple eyes (ocelli) in a characteristic depression between their two large compound eyes. The mystax helps protect the head and face when the fly encounters prey bent on defense. The antennae are short, 3-segmented, sometimes with a bristle-like structure called an arista. The short, strong proboscis is used to stab and inject victims with saliva containing neurotoxic and proteolytic enzymes which paralyze and digest the insides; the fly then sucks the liquefied meal much like we vacuum up an ice cream soda through a straw. Many species have long, tapering abdomens, sometimes with a sword-like ovipositor. Others are fat-bodied bumble bee mimics; the effect is quite convincing. Take a close look at any insect that looks like a bumble bee if it's sitting on a leaf - chances are, it's a robber fly. (Bumble bees as a rule do not sit in one spot for more than a few seconds).
References:
1.  Fritz Geller-Grimm, "Asilidae"
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Flies of North America - Order Diptera. Flies are prevalent in virtually all habitats, with over 16,000 species in North America. Flies can be distinguished from all other insects in that they only have one pair of normal wings. The other pair has evolved into small ball-like structures called halteres. Most flies have compound eyes and mouthparts adapted for piercing, lapping or sucking fluids.
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